Efferent Fibers
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Efferent nerve fibers refer to
axon An axon (from Greek ἄξων ''áxōn'', axis), or nerve fiber (or nerve fibre: see spelling differences), is a long, slender projection of a nerve cell, or neuron, in vertebrates, that typically conducts electrical impulses known as action p ...
al projections that ''exit'' a particular region; as opposed to afferent projections that ''arrive'' at the region. These terms have a slightly different meaning in the context of the
peripheral nervous system The peripheral nervous system (PNS) is one of two components that make up the nervous system of bilateral animals, with the other part being the central nervous system (CNS). The PNS consists of nerves and ganglia, which lie outside the brain ...
(PNS) and
central nervous system The central nervous system (CNS) is the part of the nervous system consisting primarily of the brain and spinal cord. The CNS is so named because the brain integrates the received information and coordinates and influences the activity of all p ...
(CNS). The efferent fiber is a long process projecting far from the neuron's body that carries
nerve impulses An action potential occurs when the membrane potential of a specific cell location rapidly rises and falls. This depolarization then causes adjacent locations to similarly depolarize. Action potentials occur in several types of animal cells, ...
away from the
central nervous system The central nervous system (CNS) is the part of the nervous system consisting primarily of the brain and spinal cord. The CNS is so named because the brain integrates the received information and coordinates and influences the activity of all p ...
toward the peripheral effector organs (mainly muscles and glands). A bundle of these fibers is called an efferent nerve (if it connects to muscles, then it is a
motor nerve A motor nerve is a nerve that transmits motor signals from the central nervous system (CNS) to the muscles of the body. This is different from the motor neuron, which includes a cell body and branching of dendrites, while the nerve is made up of ...
). The opposite direction of neural activity is afferent conduction, which carries impulses by way of the afferent nerve fibers of
sensory neuron Sensory neurons, also known as afferent neurons, are neurons in the nervous system, that convert a specific type of stimulus, via their receptors, into action potentials or graded potentials. This process is called sensory transduction. The cel ...
s. In the nervous system there is a "closed loop" system of sensation, decision, and reactions. This process is carried out through the activity of sensory neurons, interneurons, and motor neurons. In the CNS, afferent and efferent projections can be from the perspective of any given brain region. That is, each brain region has its own unique set of afferent and efferent projections. In the context of a given brain region, afferents are arriving fibers while efferents are exiting fibers.


Structure


Motor nerve

The efferent nerve fibers of motor neurons are involved in muscle control, both skeletal and smooth muscle. The
cell Cell most often refers to: * Cell (biology), the functional basic unit of life Cell may also refer to: Locations * Monastic cell, a small room, hut, or cave in which a religious recluse lives, alternatively the small precursor of a monastery ...
body Body may refer to: In science * Physical body, an object in physics that represents a large amount, has mass or takes up space * Body (biology), the physical material of an organism * Body plan, the physical features shared by a group of anima ...
of the motor neuron is connected to a single, long
axon An axon (from Greek ἄξων ''áxōn'', axis), or nerve fiber (or nerve fibre: see spelling differences), is a long, slender projection of a nerve cell, or neuron, in vertebrates, that typically conducts electrical impulses known as action p ...
and several shorter
dendrite Dendrites (from Greek δένδρον ''déndron'', "tree"), also dendrons, are branched protoplasmic extensions of a nerve cell that propagate the electrochemical stimulation received from other neural cells to the cell body, or soma, of the ...
s projecting out of the cell body itself. This axon then forms a neuromuscular junction with the effectors. The cell body of the motor neuron is
satellite A satellite or artificial satellite is an object intentionally placed into orbit in outer space. Except for passive satellites, most satellites have an electricity generation system for equipment on board, such as solar panels or radioi ...
-shaped. The motor neuron is present in the
grey matter Grey matter is a major component of the central nervous system, consisting of neuronal cell bodies, neuropil (dendrites and unmyelinated axons), glial cells (astrocytes and oligodendrocytes), synapses, and capillaries. Grey matter is distingu ...
of the
spinal cord The spinal cord is a long, thin, tubular structure made up of nervous tissue, which extends from the medulla oblongata in the brainstem to the lumbar region of the vertebral column (backbone). The backbone encloses the central canal of the sp ...
and medulla oblongata, and forms an electrochemical pathway to the effector organ or muscle. Besides motor nerves, there are efferent sensory nerves that often serve to adjust the sensitivity of the signal relayed by the afferent sensory nerve.


Types

There are three types of efferent fibers:
general somatic efferent fibers The general (spinal) somatic efferent neurons (GSE, somatomotor, or somatic motor fibers), arise from motor neuron cell bodies in the ventral horns of the gray matter within the spinal cord. They exit the spinal cord through the ventral roots, car ...
(GSE),
general visceral efferent fibers General visceral efferent fibers (GVE) or visceral efferents or autonomic efferents, are the efferent nerve fibers of the autonomic nervous system (also known as the ''visceral efferent nervous system'' that provide motor innervation to smooth mu ...
(GVE) and special visceral efferent fibers (SVE). Subtypes of general somatic efferent fibers include: alpha motor neurons (α) – these target
extrafusal muscle fiber Extrafusal muscle fibers are the standard skeletal muscle fibers that are innervated by alpha motor neurons and generate tension by contracting, thereby allowing for skeletal movement. They make up the large mass of skeletal striated muscle t ...
s, and
gamma motor neuron A gamma motor neuron (γ motor neuron), also called gamma motoneuron, or fusimotor neuron, is a type of lower motor neuron that takes part in the process of muscle contraction, and represents about 30% of ( Aγ) fibers going to the muscle. Like ...
s (γ) that target
intrafusal muscle fiber Intrafusal muscle fibers are skeletal muscle fibers that serve as specialized sensory organs ( proprioceptors). They detect the amount and rate of change in length of a muscle.Casagrand, Janet (2008) ''Action and Movement: Spinal Control o ...
s.
Beta motor neuron Beta motor neurons (β motor neurons), also called beta motoneurons, are a kind of lower motor neuron, along with alpha motor neurons and gamma motor neurons. Beta motor neurons innervate intrafusal fibers of muscle spindles with collaterals to e ...
s target both types of muscle fiber and there are two types known as ''static'' and ''dynamic''.


Etymology and mnemonics

Both afferent and efferent come from French, evolved from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
(the basis of many terms in medicine and biology) for the terms, respectively, ''ad ferens'' (Latin verb ''ferre'': carry), meaning ''carrying into'', and ''ex ferens'', meaning ''carrying away'' (''ad'' literally means ''to'', and ''e'' = ''ex'' means ''from''). ''Ad'' and ''ex'' give an easy
mnemonic device A mnemonic ( ) device, or memory device, is any learning technique that aids information retention or retrieval (remembering) in the human memory for better understanding. Mnemonics make use of elaborative encoding, retrieval cues, and imagery ...
for remembering the relationship between ''afferent'' and ''efferent'': ''afferent connection arrives'' and an ''efferent connection exits''. Afferent and efferent are connected to affect and effect through their common Latin roots: Afferent nerves affect the subject, whereas efferent nerves allow the subject to effect change.


References

{{Authority control Peripheral nervous system Medical mnemonics